The most common and most likely answer to this crossword clue is the 15 letter word TARSOMETATARSUS
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The cryptic/wordplay part
In cryptic crossword there's often an wordplay/cryptic part as well.This part involves a more complex hint, such as an anagram, homophone, hidden word, etc.
Sadly, We don't have an explanation for this specific crossword clue yet.
What does TARSOMETATARSUS mean?
TARSOMETATARSUS noun- The large bone next the foot in the leg of a bird. It is formed by the union of the distal part of the tarsus with the metatarsus.