Cryptic crossword clue
Particle requiring question mark? Millions lost 5

The most common and most likely answer to this crossword clue is the 5 letter word QUARK

I've seen this crossword clue in Financial Times
Q
U
A
R
K

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The straight part
We know that "Particle" is the straight/definition part of the clue. The straight part is often a synonym or definition, in this case particle = quark.
The cryptic/wordplay part
In cryptic crossword there's often an wordplay/cryptic part as well.
This part involves a more complex hint, such as an anagram, homophone, hidden word, etc.
Sadly, We don't have an explanation for this specific crossword clue yet.

What does QUARK mean?

QUARK noun
  1. In the Standard Model, an elementary subatomic particle that forms matter. They combine to form hadrons, such as protons and neutrons.
  2. (X Window System) An integer that uniquely identifies a text string.
QUARK noun
  1. A soft creamy cheese, eaten throughout northern, central, and eastern Europe, very similar to cottage cheese except that it is usually not made with rennet.
QUARK noun
  1. (Falkland Islands) The black-crowned night heron, Nycticorax nycticorax.

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