Cryptic crossword clue
Rudimentary toes, say, all but the first inverted in flesh of calf 9

The most common and most likely answer to this crossword clue is the 9 letter word VESTIGIAL

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The cryptic/wordplay part
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This part involves a more complex hint, such as an anagram, homophone, hidden word, etc.
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What does VESTIGIAL mean?

VESTIGIAL noun
  1. A small, degenerate, or imperfectly developed part or organ which has been more fully developed in some past generation.
VESTIGIAL adjective
  1. Of or pertaining to a vestige or remnant; like a trace from the past.
  2. Not fully developed in mature animals.

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